Kimberley Jane Hockings. Published 29 articles in journals.

Personal identification

Full name
Kimberley Jane Hockings

Citation names

  • Hockings, Kimberley

Author identifiers

Ciência ID


Language Speaking Reading Writing Listening Peer-review
English Advanced (C1) Advanced (C1) Advanced (C1) Advanced (C1)
French Intermediate (B1) Advanced (C1) Intermediate (B1) Intermediate (B1)
Spanish; Castilian Beginner (A1) Beginner (A1) Beginner (A1) Beginner (A1)
German Beginner (A1) Beginner (A1) Beginner (A1) Beginner (A1)
Portuguese Intermediate (B1) Intermediate (B1) Intermediate (B1) Intermediate (B1)
Degree Classification
PhD programme (Doutoramento)
University of Stirling, United Kingdom
"Human-chimpanzee coexistence at Bossou, the Republic of Guinea: A chimpanzee perspective" (THESIS/DISSERTATION)
PhD awarded
Zoology (Bacharelato)
University of Liverpool Faculty of Health and Life Sciences, United Kingdom
"Allogrooming and proximity in chimpanzees during the feeding period: association with conflict" (THESIS/DISSERTATION)
BSc with honours. Upper 2:1 degree


Host institution
2018/08/01 - Current Senior Lecturer University of Exeter Centre for Ecology and Conservation, United Kingdom
2015/03/01 - Current FCT INVESTIGADOR/RESEARCH FELLOW - To continue to develop Primatology in Portugal, to conduct world-class scientific research, to supervise and teach Master's and PhD students, and to contribute to ac Centro em Rede de Investigação em Antropologia, Portugal
2008 - Current Research Group Member Centro em Rede de Investigação em Antropologia, Portugal
2013/09/01 - 2014/08/01 POSTDOCTORAL RESEARCH FELLOW - To conduct research at an International/world-leading standard; To develop independent & collaborative grant applications; To teach on Undergrad & Masters courses; To s Oxford Brookes University, United Kingdom
2012/08/01 - 2013/08/30 FIELD SCHOOL INSTRUCTOR for 1 month in 2012/2013, Kenya - To teach key concepts in primatology & wildlife conservation; To organise interactive workshops and role-playing exercises; To supervise resea Rutgers University Department of Anthropology, United States
2011/03/01 - 2013/08/01 VISITING RESEARCH FELLOW - To conduct research of an International & world-leading standard Oxford Brookes University, United Kingdom
2008/03/01 - 2013/08/01 POSTDOCTORAL RESEARCHER - To conduct research at an International standard; To teach on BA & PhD programmes; To write grants to International & Portuguese funding bodies; To organise symposia & confer Universidade Nova de Lisboa, Portugal
2008/03/01 - 2011/02/01 HONORARY RESEARCH FELLOW - To conduct research of an International & world-leading standard University of Stirling, United Kingdom
2010/02/01 - 2010/04/01 VISITING POSTDOCTORAL RESEARCHER- To develop research collaborations between ethologists, geneticists, archaeologists & evolutionary psychologists; To produce research articles; To organise a cross-di Kyoto Daigaku, Japan


Designation Funders
2019/04 - 2022/03 Promoting public health in a biodiverse agroforest landscape in Guinea-Bissau
United Kingdom Department for Environment Food and Rural Affairs


Journal article
  1. Zoe M. Satsias; Matthew J. Silk; Kimberley J. Hockings; Marie Cibot; Jacqueline Rohen; Matthew R. McLennan. "Sex-specific responses to anthropogenic risk shape wild chimpanzee social networks in a human-impacted landscape". Animal Behaviour (2022): https://doi.org/10.1016/j.anbehav.2022.01.016.
  2. Max Bain; Arsha Nagrani; Daniel Schofield; Sophie Berdugo; Joana Bessa; Jake Owen; Kimberley J. Hockings; et al. "Automated audiovisual behavior recognition in wild primates". Science Advances 7 46 (2021): https://doi.org/10.1126/sciadv.abi4883.
  3. Balint Andrasi; Jochen A.G. Jaeger; Stefanie Heinicke; Kristian Metcalfe; Kimberley J. Hockings. "Quantifying the road-effect zone for a critically endangered primate". Conservation Letters (2021): https://doi.org/10.1111/conl.12839.
  4. Kimberley J. Hockings; Benjamin Mubemba; Charlotte Avanzi; Kamilla Pleh; Ariane Düx; Elena Bersacola; Joana Bessa; et al. "Leprosy in wild chimpanzees". Nature (2021): https://doi.org/10.1038/s41586-021-03968-4.
  5. Ruth H. Thurstan; Kimberley J. Hockings; Johanna S. U. Hedlund; Elena Bersacola; Claire Collins; Regan Early; Yunsiska Ermiasi; et al. "Envisioning a resilient future for biodiversity conservation in the wake of the COVID-19 pandemic". People and Nature (2021): https://doi.org/10.1002/pan3.10262.
  6. Hockings, Kimberley. "Chimpanzees balance resources and risk in an anthropogenic landscape of fear". Scientific Reports (2021): https://doi.org/10.1038/s41598-021-83852-3.
    In press • 10.1038/s41598-021-83852-3
  7. Heinicke, Stefanie; Ordaz-Németh, Isabel; Junker, Jessica; Bachmann, Mona E.; Marrocoli, Sergio; Wessling, Erin G.; Byler, Dirck; et al. "Open-access platform to synthesize knowledge of ape conservation across sites". American Journal of Primatology 83 1 (2020): http://dx.doi.org/10.1002/ajp.23213.
  8. McLennan, Matthew R.; Hintz, Benjamin; Kiiza, Vicent; Rohen, Jacqueline; Lorenti, Georgia A.; Hockings, Kimberley J.. "Surviving at the extreme: Chimpanzee ranging is not restricted in a deforested human-dominated landscape in Uganda". African Journal of Ecology (2020): http://dx.doi.org/10.1111/aje.12803.
  9. Hockings, Kimberley. "Extensive overlap in the selection of wild fruits by chimpanzees and humans: Implications for the management of complex social-ecological systems". Frontiers in Ecology and the Environment (2020): https://www.frontiersin.org/articles/10.3389/fevo.2020.00123/abstract.
    Open access • Accepted
  10. Wilson F. Vieira; Chris Kerry; Kimberley J. Hockings. "A comparison of methods to determine chimpanzee home-range size in a forest–farm mosaic at Madina in Cantanhez National Park, Guinea-Bissau". Primates 60 4 (2019): 355-365. https://doi.org/10.1007/s10329-019-00724-1.
  11. Hannah E. Parathian; Matthew R. McLennan; Catherine M. Hill; Amélia Frazão-Moreira; Kimberley J. Hockings. "Breaking Through Disciplinary Barriers: Human–Wildlife Interactions and Multispecies Ethnography". International Journal of Primatology 39 5 (2018): 749-775. https://doi.org/10.1007/s10764-018-0027-9.
  12. Kimberley J. Hockings. "An Introduction to Primate Conservation, edited by Serge A. Wich and Andrew J. Marshall: a review". Primates 58 1 (2017): 259-260. https://doi.org/10.1007%2Fs10329-016-0589-0.
  13. Hockings, K.J.; Yamakoshi, G.; Matsuzawa, T.. "Dispersal of a Human-Cultivated Crop by Wild Chimpanzees (Pan troglodytes verus) in a Forest–Farm Matrix". International Journal of Primatology 38 2 (2017): 172-193. http://www.scopus.com/inward/record.url?eid=2-s2.0-84994067266&partnerID=MN8TOARS.
  14. McLennan, M.R.; Spagnoletti, N.; Hockings, K.J.. "The Implications of Primate Behavioral Flexibility for Sustainable Human–Primate Coexistence in Anthropogenic Habitats". International Journal of Primatology 38 2 (2017): 105-121. http://www.scopus.com/inward/record.url?eid=2-s2.0-85018255222&partnerID=MN8TOARS.
  15. Fernandes, M.; Frazão-Moreira, A.; Hockings, K.J.; Alves-Cardoso, F.. "Across disciplinary boundaries: Remembering cláudia sousa | Para além das fronteiras disciplinares: Lembrando cláudia sousa". Etnografica 20 3 (2016): 633-640. http://www.scopus.com/inward/record.url?eid=2-s2.0-85000869331&partnerID=MN8TOARS.
  16. Hockings, K.J.. "Chimpanzee behavioural flexibility and the sustainability of human-chimpanzee interactions at cantanhez national park, guinea-bissau | Flexibilidade comportamental dos chimpanzés e a sustentabilidade das interações entre humanos e chimpanzés no parque nacional de cantanhez, guiné-bissau". Etnografica 20 3 (2016): 659-662. http://www.scopus.com/inward/record.url?eid=2-s2.0-85000394603&partnerID=MN8TOARS.
  17. Cannon, T.H.; Heistermann, M.; Hankison, S.J.; Hockings, K.J.; McLennan, M.R.. "Tailored Enrichment Strategies and Stereotypic Behavior in Captive Individually Housed Macaques (Macaca spp.)". Journal of Applied Animal Welfare Science 19 2 (2016): 171-182. http://www.scopus.com/inward/record.url?eid=2-s2.0-84961206682&partnerID=MN8TOARS.
  18. Fujisawa, M.; Hockings, K.J.; Soumah, A.G.; Matsuzawa, T.. "Placentophagy in wild chimpanzees (Pan troglodytes verus) at bossou, guinea". Primates 57 2 (2016): 175-180. http://www.scopus.com/inward/record.url?eid=2-s2.0-84954318569&partnerID=MN8TOARS.
  19. Kimberley J. Hockings; Nicola Bryson-Morrison; Susana Carvalho; Michiko Fujisawa; Tatyana Humle; William C. McGrew; Miho Nakamura; et al. "Tools to tipple: ethanol ingestion by wild chimpanzees using leaf-sponges". Royal Society Open Science 2 6 (2015): 150150-150150. https://doi.org/10.1098/rsos.150150.
  20. Kimberley J. Hockings; Matthew R. McLennan; Susana Carvalho; Marc Ancrenaz; René Bobe; Richard W. Byrne; Robin I.M. Dunbar; et al. "Apes in the Anthropocene: flexibility and survival". Trends in Ecology & Evolution 30 4 (2015): 215-222. http://dx.doi.org/10.1016/j.tree.2015.02.002.
  21. Joana Bessa; Cláudia Sousa; Kimberley J. Hockings. "Feeding ecology of chimpanzees ( Pan troglodytes verus ) inhabiting a forest-mangrove-savanna-agricultural matrix at Caiquene-Cadique, Cantanhez National Park, Guinea-Bissau". Am. J. Primatol. (2015): n/a-n/a. http://dx.doi.org/10.1002/ajp.22388.
  22. Hockings, Kimberley. "Wild chimpanzees show group differences in selection of agricultural crops". Scientific Reports (2014):
  23. Kimberley Hockings. "Fear beyond predators". Science (2014):
  24. Hockings, K.J.; McLennan, M.R.; Hill, C.. "Fear beyond predators". Science 344 6187 (2014): http://www.scopus.com/inward/record.url?eid=2-s2.0-84901639724&partnerID=MN8TOARS.
  25. Hockings, K.J.; Sousa, C.. "Human-chimpanzee sympatry and interactions in Cantanhez National Park, guinea-bissau: Current research and future directions". Primate Conservation 26 1 (2013): 57-65. http://www.scopus.com/inward/record.url?eid=2-s2.0-84875196334&partnerID=MN8TOARS.
  26. Hockings, K.J.; McLennan, M.R.. "From forest to farm: Systematic review of cultivar feeding by chimpanzees - management implications for wildlife in anthropogenic landscapes". PLoS ONE 7 4 (2012): http://www.scopus.com/inward/record.url?eid=2-s2.0-84859560845&partnerID=MN8TOARS.
  27. Carvalho, S.; Biro, D.; Cunha, E.; Hockings, K.; McGrew, W.C.; Richmond, B.G.; Matsuzawa, T.. "Chimpanzee carrying behaviour and the origins of human bipedality". Current Biology 22 6 (2012): http://www.scopus.com/inward/record.url?eid=2-s2.0-84858674149&partnerID=MN8TOARS.
  28. Hockings, K.J.; Anderson, J.R.; Matsuzawa, T.. "Socioecological adaptations by chimpanzees, Pan troglodytes verus, inhabiting an anthropogenically impacted habitat". Animal Behaviour 83 3 (2012): 801-810. http://www.scopus.com/inward/record.url?eid=2-s2.0-84862776709&partnerID=MN8TOARS.
  29. Little, A.C.; Hockings, K.J.; Apicella, C.L.; Sousa, C.. "Mixed-ethnicity face shape and attractiveness in humans". Perception 41 12 (2012): 1486-1496. http://www.scopus.com/inward/record.url?eid=2-s2.0-84874202480&partnerID=MN8TOARS.
  30. Hockings, K.J.; Sousa, C.. "Differential utilization of cashew-a low-conflict crop-by sympatric humans and chimpanzees". ORYX 46 3 (2012): 375-381. http://www.scopus.com/inward/record.url?eid=2-s2.0-84863912790&partnerID=MN8TOARS.
  31. Hockings, K.J.; Humle, T.; Carvalho, S.; Matsuzawa, T.. "Chimpanzee interactions with nonhuman species in an anthropogenic habitat". Behaviour 149 3-4 (2012): 299-324. http://www.scopus.com/inward/record.url?eid=2-s2.0-84861675783&partnerID=MN8TOARS.
  32. Hockings, K.J.; Yamakoshi, G.; Kabasawa, A.; Matsuzawa, T.. "Attacks on local persons by chimpanzees in Bossou, Republic of Guinea: Long-term perspectives". American Journal of Primatology 72 10 (2010): 887-896. http://www.scopus.com/inward/record.url?eid=2-s2.0-77957101109&partnerID=MN8TOARS.
  33. Hockings, K.J.; Anderson, J.R.; Matsuzawa, T.. "Flexible feeding on cultivated underground storage organs by rainforest-dwelling chimpanzees at Bossou, West Africa". Journal of Human Evolution 58 3 (2010): 227-233. http://www.scopus.com/inward/record.url?eid=2-s2.0-76949102586&partnerID=MN8TOARS.
  34. Fuentes, A.; Hockings, K.J.. "The ethnoprimatological approach in primatology". American Journal of Primatology 72 10 (2010): 841-847. http://www.scopus.com/inward/record.url?eid=2-s2.0-77957077080&partnerID=MN8TOARS.
  35. Hockings, K.J.; Anderson, J.R.; Matsuzawa, T.. "Use of wild and cultivated foods by Chimpanzees at Bossou, Republic of Guinea: Feeding dynamics in a human-influenced environment". American Journal of Primatology 71 8 (2009): 636-646. http://www.scopus.com/inward/record.url?eid=2-s2.0-67849121527&partnerID=MN8TOARS.
  36. Hockings, K.J.. "Living at the interface: Human-chimpanzee competition, coexistence and conflict in Africa". Interaction Studies 10 2 (2009): 183-205. http://www.scopus.com/inward/record.url?eid=2-s2.0-70349324176&partnerID=MN8TOARS.
  37. Hockings, K.J.; Humle, T.; Anderson, J.R.; Biro, D.; Sousa, C.; Ohashi, G.; Matsuzawa, T.. "Chimpanzees share forbidden fruit". PLoS ONE 2 9 (2007): http://www.scopus.com/inward/record.url?eid=2-s2.0-41149086654&partnerID=MN8TOARS.
  38. Hockings, K.J.; Anderson, J.R.; Matsuzawa, T.. "Road crossing in chimpanzees: A risky business". Current Biology 16 17 (2006): http://www.scopus.com/inward/record.url?eid=2-s2.0-33750615073&partnerID=MN8TOARS.
Working paper
  1. Hockings, Kimberley. 2020. "Regional Action Plan for the Conservation of Western Chimpanzees (Pan troglodytes verus) 2020–2030". Working Paper. doi.org/10.2305/IUCN.CH.2020.SSC-RAP.2.en.


Other output
  1. Leprosy in wild chimpanzees. 2020. Kimberley J. Hockings; Benjamin Mubemba; Charlotte Avanzi; Kamilla Pleh; Ariane Düx; Elena Bersacola; Joana Bessa; et al. https://doi.org/10.1101/2020.11.10.374371.
  2. The Aggressive Apes? Causes and Contexts of Great Ape Attacks on Local Persons. 2015. Matthew R. McLennan; Kimberley J. Hockings. http://dx.doi.org/10.1007/978-3-319-22246-2_18.

Other distinction

2016 Conservation Workshop grant
2016 Santander Internationalisation Award for Scientific Production - €625
2013 Santander Internationalisation Award for Scientific Production - €1250
2010 Conference grant - €2000
2010 Research Equipment - US$2500
2009 Research grant - US$4500
2008 Post-doctoral research grant and extension (2008 – 2013); International travel grants (2008 – present) - €4600; Conference grants (2008 – present) - €5450.
2004 Fieldwork costs (2004 – present, distributed by T Matsuzawa) - £15000.
2003 Full PhD studentship (2003-2007), Research & conference grants (2004-2006) - £1940