Xavier Santos. Completed the Doktor (PhD) in Biologia Animal. Zoologia in 2000/09/22 by Universitat de Barcelona Facultat de Biologia and Bachelor in Zoology in 1984/10/04 by Universitat de Barcelona Facultat de Biologia. Is Teacher in a Public Secondary School in Generalitat de Catalunya Departament d'Ensenyament. Published 102 articles in journals. Has received 1 awards and/or honors.

Identificação pessoal

Nome completo
Xavier Santos

Nomes de citação

  • Santos, Xavier

Identificadores de autor

Ciência ID

Endereços de correio eletrónico

  • xsantossantiro@gmail.com (Profissional)

Domínios de atuação

  • Ciências Naturais - Ciências Biológicas - Ecologia
  • Ciências Naturais - Ciências Biológicas - Ecologia


Idioma Conversação Leitura Escrita Compreensão Peer-review
Espanhol; Castelhano Utilizador proficiente (C1) Utilizador proficiente (C1) Utilizador proficiente (C1) Utilizador proficiente (C1)
Inglês Utilizador independente (B1) Utilizador proficiente (C1) Utilizador proficiente (C1) Utilizador independente (B1)
Francês Utilizador independente (B1) Utilizador proficiente (C1) Utilizador elementar (A1) Utilizador independente (B1)
Catalão Utilizador proficiente (C1) Utilizador proficiente (C1) Utilizador proficiente (C1) Utilizador proficiente (C1)
Português Utilizador independente (B1) Utilizador independente (B1) Utilizador elementar (A1) Utilizador independente (B1)
Grau Classificação
Biologia Animal. Zoologia (Doktor (PhD))
Universitat de Barcelona Facultat de Biologia, Espanha
"Ecology of the viperine snake Natrix maura (Linnaeus, 1758) in the Ebro delta" (TESE/DISSERTAÇÃO)
Excellent Cum laude
Zoology (Bachelor)
Universitat de Barcelona Facultat de Biologia, Espanha
Percurso profissional


Categoria Profissional
Instituição de acolhimento
2011/09/01 - 2016/08/31 Pós-doutorado (Investigação) Universidade do Porto Centro de Investigação em Biodiversidade e Recursos Genéticos, Portugal
Universidade do Porto Centro de Investigação em Biodiversidade e Recursos Genéticos, Portugal
2010/03 - 2010/07/31 Pós-doutorado (Investigação) Centre d'Ecologie Fonctionnelle et Evolutive, França
Centre d'Ecologie Fonctionnelle et Evolutive, França
2006/09/01 - 2008/08/31 Investigador (Investigação) Diputació de Barcelona, Espanha
Parc Natural de Sant Llorenç del Munt i l'Obac, Espanha
2003/09/01 - 2005/08/31 Pós-doutorado (Investigação) Universidad de Granada, Espanha
Universidad de Granada Facultad de Ciencias, Espanha


Categoria Profissional
Instituição de acolhimento
2016/09/01 - 2022/08/31 Teacher in a Public Secondary School Generalitat de Catalunya Departament d'Ensenyament, Espanha
Institut Les Corts, Espanha


Artigo em revista
  1. Roger Puig-Gironès; Xavier Santos; Vicenç Bros. "Long-interval effects of wildfires on the functional diversity of land snails". Science of The Total Environment (2023): https://doi.org/10.1016/j.scitotenv.2023.162677.
  2. Roger Puig-Gironès; Xavier Santos; Vicenç Bros. "Temporal differences in snail diversity responses to wildfires and salvage logging". Environmental Conservation (2023): https://doi.org/10.1017/S0376892922000443.
  3. Xavier Santos; Juan M. Pleguezuelos; Brahim Chergui; Philippe Geniez; Marc Cheylan. "Citizen-science data shows long-term decline of snakes in southwestern Europe". Biodiversity and Conservation 31 5-6 (2022): 1609-1625. https://doi.org/10.1007/s10531-022-02415-8.
  4. Xavier Santos; Josabel Belliure; João F. Gonçalves; Juli G. Pausas. "Resilience of reptiles to megafires". Ecological Applications (2022): https://doi.org/10.1002/eap.2518.
  5. Fernando Martínez-Freiría; Inês Freitas; Guillermo Velo-Antón; Nahla Lucchini; Soumia Fahd; Said Larbes; Juan M. Pleguezuelos; Xavier Santos; José C. Brito. "Integrative taxonomy reveals two species and intraspecific differentiation in the Vipera latastei–monticola complex". Journal of Zoological Systematics and Evolutionary Research (2021): https://doi.org/10.1111/jzs.12534.
  6. Alberto Muñoz; Ángel M. Felicísimo; Xavier Santos. "Analysing How Pre-Fire Habitat Legacy and Post-Fire Management Influence the Resilience of Reptiles to Fire". Forests (2021): https://doi.org/10.3390/f12111487.
  7. Álvarez-Ruiz, Lola; Belliure, Josabel; Santos, Xavier; Pausas, Juli G.. "Fire reduces parasite load in a Mediterranean lizard". Proceedings of the Royal Society B: Biological Sciences 288 1954 (2021): 20211230. http://dx.doi.org/10.1098/rspb.2021.1230.
    Publicado • 10.1098/rspb.2021.1230
  8. Català, Cristina; Bros, Vicenç; Castelltort, Xavier; Santos, Xavier; Pascual, Marta. "Deep genetic structure at a small spatial scale in the endangered land snail Xerocrassa montserratensis". Scientific Reports 11 1 (2021): http://dx.doi.org/10.1038/s41598-021-87741-7.
    Acesso aberto • Publicado • 10.1038/s41598-021-87741-7
  9. Roca, Vicente; Belliure, Josabel; Santos, Xavier; Pausas, Juli G.. "New Reptile Hosts for Helminth Parasites in a Mediterranean Region". Journal of Herpetology 54 2 (2020): 268. http://dx.doi.org/10.1670/18-133.
  10. Graciá, E.; Rodríguez-Caro, R.C.; Ferrández, M.; Martínez-Silvestre, A.; Pérez-Ibarra, I.; Amahjour, R.; Aranda, C.; et al. "From troubles to solutions: Conservation of mediterranean tortoises under global change". Basic and Applied Herpetology 34 (2020): 5-16. http://www.scopus.com/inward/record.url?eid=2-s2.0-85098569536&partnerID=MN8TOARS.
  11. Chergui, B.; Pleguezuelos, J.M.; Fahd, S.; Santos, X.. "Modelling functional response of reptiles to fire in two Mediterranean forest types". Science of the Total Environment 732 (2020): http://www.scopus.com/inward/record.url?eid=2-s2.0-85084657964&partnerID=MN8TOARS.
  12. Freitas, I.; Ursenbacher, S.; Mebert, K.; Zinenko, O.; Schweiger, S.; Wüster, W.; Brito, J.C.; et al. "Evaluating taxonomic inflation: Towards evidence-based species delimitation in Eurasian vipers (Serpentes: Viperinae)". Amphibia Reptilia 41 3 (2020): 285-311. http://www.scopus.com/inward/record.url?eid=2-s2.0-85085036041&partnerID=MN8TOARS.
  13. Jiménez-Albarral, J.J.; Pleguezuelos, J.M.; Santos, X.. "Positive effect of catastrophic winds on reptile community recovery in pine plantations". Basic and Applied Ecology 46 (2020): 1-9. http://www.scopus.com/inward/record.url?eid=2-s2.0-85085970294&partnerID=MN8TOARS.
  14. Fuentes, I.; Márquez-Ferrando, R.; Pleguezuelos, J.M.; Sanpera, C.; Santos, X.. "Long-term trace element assessment after a mine spill: Pollution persistence and bioaccumulation in the trophic web". Environmental Pollution 267 (2020): http://www.scopus.com/inward/record.url?eid=2-s2.0-85089883211&partnerID=MN8TOARS.
  15. Diana Ferreira; Catarina Pinho; José Carlos Brito; Xavier Santos. "Increase of genetic diversity indicates ecological opportunities in recurrent-fire landscapes for wall lizards". Scientific Reports 9 1 (2019): https://doi.org/10.1038/s41598-019-41729-6.
  16. Brahim Chergui; Teresa Morán-López; Roberto C. Rodríguez-Caro; Eva Graciá; Soumia Fahd; Xavier Santos. "Population density of the spur-thighed tortoise Testudo graeca declines after fire in north-western Africa". PLOS ONE (2019): https://doi.org/10.1371/journal.pone.0220969.
  17. Alberto Muñoz; Ángel M. Felicísimo; Xavier Santos. "Assessing the resistance of a breeding amphibian community to a large wildfire". Acta Oecologica 99 (2019): 103439-103439. https://doi.org/10.1016/j.actao.2019.06.002.
  18. Karin Tamar; Marc Simó-Riudalbas; Joan Garcia-Porta; Xavier Santos; Gustavo Llorente; Raquel Vasconcelos; Salvador Carranza. "An integrative study of island diversification: Insights from the endemic Haemodracon geckos of the Socotra Archipelago". Molecular Phylogenetics and Evolution 133 (2019): 166-175. https://doi.org/10.1016/j.ympev.2019.01.009.
  19. Brahim Chergui; Soumia Fahd; Xavier Santos. "Are reptile responses to fire shaped by forest type and vegetation structure? Insights from the Mediterranean basin". Forest Ecology and Management 437 (2019): 340-347. https://doi.org/10.1016/j.foreco.2019.01.046.
  20. Xavier Santos; Neftalí Sillero; Françoise Poitevin; Marc Cheylan. "Realized niche modelling uncovers contrasting responses to fire according to species-specific biogeographical affinities of amphibian and reptile species". Biological Journal of the Linnean Society 126 1 (2019): 55-67. https://doi.org/10.1093/biolinnean/bly173.
  21. Lino, S.; Sillero, N.; Torres, J.; Santos, X.; Álvares, F.. "The role of fire on wolf distribution and breeding-site selection: Insights from a generalist carnivore occurring in a fire-prone landscape". Landscape and Urban Planning 183 (2019): 111-121. http://www.scopus.com/inward/record.url?eid=2-s2.0-85058159459&partnerID=MN8TOARS.
  22. Eduardo MATEOS; Marta GOULA; Teresa SAURAS; Xavier SANTOS. "Habitat structure and host plant specialization drive taxonomic andfunctional composition of Heteroptera in postfire successional habitats". TURKISH JOURNAL OF ZOOLOGY 42 4 (2018): 449-463. https://doi.org/10.3906/zoo-1710-21.
  23. Tiago Pinto; Bruno Moreira; Helena Freitas; Xavier Santos. "The role of fire history, land-use, and vegetation structure on the response of Mediterranean lizards to fire". Forest Ecology and Management 419-420 (2018): 139-145. https://doi.org/10.1016/j.foreco.2018.03.029.
  24. Xavier Santos; Sandra Navarro; João Carlos Campos; Carola Sanpera; José Carlos Brito. "Stable isotopes uncover trophic ecology of the West African crocodile (Crocodylus suchus)". Journal of Arid Environments 148 (2018): 6-13. https://doi.org/10.1016/j.jaridenv.2017.09.008.
  25. Chergui, B.; Fahd, S.; Santos, X.; Pausas, J.G.. "Socioeconomic Factors Drive Fire-Regime Variability in the Mediterranean Basin". Ecosystems 21 4 (2018): 619-628. http://www.scopus.com/inward/record.url?eid=2-s2.0-85028310951&partnerID=MN8TOARS.
  26. Chergui, B.; Fahd, S.; Santos, X.. "Quercus suber forest and Pinus plantations show different post-fire resilience in Mediterranean north-western Africa". Annals of Forest Science 75 2 (2018): http://www.scopus.com/inward/record.url?eid=2-s2.0-85048373791&partnerID=MN8TOARS.
  27. Ferreira, D.; Brito, J.C.; Santos, X.. "Long-interval monitoring reveals changes in the structure of a reptile community in a biogeographic transition zone". Basic and Applied Herpetology 32 (2018): 41-55. http://www.scopus.com/inward/record.url?eid=2-s2.0-85044525352&partnerID=MN8TOARS.
  28. Santos, X.; Azor, J.S.; Cortés, S.; Rodríguez, E.; Larios, J.; Pleguezuelos, J.M.. "Ecological significance of dorsal polymorphism in a Batesian mimic snake". Current Zoology 64 6 (2018): 745-753. http://www.scopus.com/inward/record.url?eid=2-s2.0-85050961880&partnerID=MN8TOARS.
  29. Sancho, V.; Vasconcelos, R.; Llorente, G.; Santos, X.. "Morphological differentiation of the complex Pristurus sokotranus (Squamata: Sphaerodactylidae) from Socotra (Yemen)". Zootaxa 4324 1 (2017): 63-84. http://www.scopus.com/inward/record.url?eid=2-s2.0-85029904871&partnerID=MN8TOARS.
  30. Badiane, A.; Matos, C.; Santos, X.. "Uncovering environmental, land-use and fire effects on the distribution of a low-dispersal species, the Hermann's tortoise Testudo hermanni". Amphibia Reptilia 38 1 (2017): 67-77. http://www.scopus.com/inward/record.url?eid=2-s2.0-85013356859&partnerID=MN8TOARS.
  31. Hudson, L.N.; Newbold, T.; Contu, S.; Hill, S.L.L.; Lysenko, I.; De Palma, A.; Phillips, H.R.P.; et al. "The database of the PREDICTS (Projecting Responses of Ecological Diversity In Changing Terrestrial Systems) project". Ecology and Evolution 7 1 (2017): 145-188. http://www.scopus.com/inward/record.url?eid=2-s2.0-85006925546&partnerID=MN8TOARS.
  32. Martínez-Freiría, F.; Pérez i de Lanuza, G.; Pimenta, A.A.; Pinto, T.; Santos, X.. "Aposematism and crypsis are not enough to explain dorsal polymorphism in the Iberian adder". Acta Oecologica 85 (2017): 165-173. http://www.scopus.com/inward/record.url?eid=2-s2.0-85033606987&partnerID=MN8TOARS.
  33. Ferreira, D.; Žagar, A.; Santos, X.. "Uncovering the rules of (Reptile) species coexistence in transition zones between bioregions". Salamandra 53 2 (2017): 193-200. http://www.scopus.com/inward/record.url?eid=2-s2.0-85019163307&partnerID=MN8TOARS.
  34. Martín, N.; Martínez, S.; Pujol-Buxó, E.; Viñolas, A.; Llorente, G.A.; Sanpera, C.; Vasconcelos, R.; Carranza, S.; Santos, X.. "Stable isotopes and diet uncover trophic-niche divergence and ecological diversification processes of endemic reptiles on Socotra Island". Zoologischer Anzeiger 267 (2017): 69-81. http://www.scopus.com/inward/record.url?eid=2-s2.0-85014035143&partnerID=MN8TOARS.
  35. Vasconcelos, R.; Rocha, S.; Santos, X.. "Sharing refuges on arid islands: Ecological and social influence on aggregation behaviour of wall geckos". PeerJ 2017 1 (2017): http://www.scopus.com/inward/record.url?eid=2-s2.0-85013187488&partnerID=MN8TOARS.
  36. Pleguezuelos, J.M.; García-Cardenete, L.; Caro, J.; Feriche, M.; Pérez-García, M.T.; Santos, X.; Sicilia, M.; Fahd, S.. "Barriers for conservation: Mitigating the impact on amphibians and reptiles by water cisterns in arid environments". Amphibia Reptilia 38 1 (2017): 113-118. http://www.scopus.com/inward/record.url?eid=2-s2.0-85013472824&partnerID=MN8TOARS.
  37. Garriga, N.; Franch, M.; Santos, X.; Montori, A.; Llorente, G.A.. "Seasonal variation in vertebrate traffic casualties and its implications for mitigation measures". Landscape and Urban Planning 157 (2017): 36-44. http://www.scopus.com/inward/record.url?eid=2-s2.0-84979536718&partnerID=MN8TOARS.
  38. Feriche, M.; Reguera, S.; Santos, X.; Mociño-Deloya, E.; Setser, K.; Pleguezuelos, J.M.. "Female Reproduction in Thamnophis scaliger: The Significance of Parturition Timing". Journal of Herpetology 50 2 (2016): 209-215. http://www.scopus.com/inward/record.url?eid=2-s2.0-84974593523&partnerID=MN8TOARS.
  39. Brito, J.C.; Tarroso, P.; Vale, C.G.; Martínez-Freiría, F.; Boratynski, Z.; Campos, J.C.; Ferreira, S.; et al. "Conservation Biogeography of the Sahara-Sahel: Additional protected areas are needed to secure unique biodiversity". Diversity and Distributions 22 4 (2016): 371-384. http://www.scopus.com/inward/record.url?eid=2-s2.0-84960118871&partnerID=MN8TOARS.
  40. Ferreira, D.; Mateus, C.; Santos, X.. "Responses of reptiles to fire in transition zones are mediated by bioregion affinity of species". Biodiversity and Conservation 25 8 (2016): 1543-1557. http://www.scopus.com/inward/record.url?eid=2-s2.0-84968572432&partnerID=MN8TOARS.
  41. Muñoz, A.; Santos, X.; Felicísimo, Á.M.. "Local-scale models reveal ecological niche variability in amphibian and reptile communities from two contrasting biogeographic regions". PeerJ 2016 10 (2016): http://www.scopus.com/inward/record.url?eid=2-s2.0-84992143213&partnerID=MN8TOARS.
  42. Vasconcelos, R.; Montero-Mendieta, S.; Simó-Riudalbas, M.; Sindaco, R.; Santos, X.; Fasola, M.; Llorente, G.; Razzetti, E.; Carranza, S.. "Unexpectedly high levels of cryptic diversity uncovered by a complete DNA barcoding of reptiles of the socotra archipelago". PLoS ONE 11 3 (2016): http://www.scopus.com/inward/record.url?eid=2-s2.0-84961958188&partnerID=MN8TOARS.
  43. Ferreira, C.C.; Santos, X.; Carretero, M.A.. "Does ecophysiology mediate reptile responses to fire regimes? Evidence from Iberian lizards". PeerJ 2016 6 (2016): http://www.scopus.com/inward/record.url?eid=2-s2.0-84977177485&partnerID=MN8TOARS.
  44. Santos, X.; Badiane, A.; Matos, C.. "Contrasts in short- and long-term responses of Mediterranean reptile species to fire and habitat structure". Oecologia 180 1 (2016): 205-216. http://www.scopus.com/inward/record.url?eid=2-s2.0-84952865114&partnerID=MN8TOARS.
  45. Bros, V.; Torre, I.; Santos, X.. "Uncovering the environmental factors that influence diversity patterns of Mediterranean terrestrial Gastropod communities: a useful tool for conservation". Ecological Research 31 1 (2016): 39-47. http://www.scopus.com/inward/record.url?eid=2-s2.0-84944711683&partnerID=MN8TOARS.
  46. Ballouard, J.-M.; Mullin, S.J.; Ajtic, R.; Brito, J.C.; ElMouden, E.H.; Erdogan, M.; Feriche, M.; et al. "Factors Influencing Schoolchildren's Responses to a Questionnaire in Wildlife Conservation Education". International Journal of Science Education 37 3 (2015): 469-483. http://www.scopus.com/inward/record.url?eid=2-s2.0-84922234444&partnerID=MN8TOARS.
  47. Martínez-Freiría, F.; Santos, X.. "Assessing the heritability of dorsal pattern shape in Vipera latastei". Amphibia Reptilia 36 3 (2015): 313-317. http://www.scopus.com/inward/record.url?eid=2-s2.0-84941058107&partnerID=MN8TOARS.
  48. Brito, J.C.; Godinho, R.; Martínez-Freiría, F.; Pleguezuelos, J.M.; Rebelo, H.; Santos, X.; Vale, C.G.; et al. "Unravelling biodiversity, evolution and threats to conservation in the sahara-sahel". Biological Reviews 89 1 (2014): 215-231. http://www.scopus.com/inward/record.url?eid=2-s2.0-84891658537&partnerID=MN8TOARS.
  49. García-Cardenete, L.; Pleguezuelos, J.M.; Brito, J.C.; Jiménez-Cazalla, F.; Pérez-García, M.T.; Santos, X.. "Water cisterns as death traps for amphibians and reptiles in arid environments". Environmental Conservation 41 4 (2014): 341-349. http://www.scopus.com/inward/record.url?eid=2-s2.0-84908874368&partnerID=MN8TOARS.
  50. Torre, I.; Bros, V.; Santos, X.. "Assessing the impact of reforestation on the diversity of Mediterranean terrestrial Gastropoda". Biodiversity and Conservation 23 10 (2014): 2579-2589. http://www.scopus.com/inward/record.url?eid=2-s2.0-84905278910&partnerID=MN8TOARS.
  51. Filippakopoulou, A.; Santos, X.; Feriche, M.; Pleguezuelos, J.M.; Llorente, G.A.. "Effect of prey availability on growth-rate trajectories of an aquatic predator, the viperine snake natrix maura | Efecto de la disponibilidad de presas en las trayectorias de crecimiento de un depredador acuático, la culebraviperina Natrix maura". Basic and Applied Herpetology 28 (2014): 35-50. http://www.scopus.com/inward/record.url?eid=2-s2.0-84928987884&partnerID=MN8TOARS.
  52. Santos, X.; Vidal-García, M.; Brito, J.C.; Fahd, S.; Llorente, G.A.; Martínez-Freiría, F.; Parellada, X.; Pleguezuelos, J.M.; Sillero, N.. "Phylogeographic and environmental correlates support the cryptic function of the zigzag pattern in a European viper". Evolutionary Ecology 28 4 (2014): 611-626. http://www.scopus.com/inward/record.url?eid=2-s2.0-84901839246&partnerID=MN8TOARS.
  53. Santos, X.; Mateos, E.; Bros, V.; Brotons, L.; De Mas, E.; Herraiz, J.A.; Herrando, S.; et al. "Is response to fire influenced by dietary specialization and mobility? A comparative study with multiple animal assemblages". PLoS ONE 9 2 (2014): http://www.scopus.com/inward/record.url?eid=2-s2.0-84895741149&partnerID=MN8TOARS.
  54. Velo-Antón, G.; Santos, X.; Sanmartín-Villar, I.; Cordero-Rivera, A.; Buckley, D.. "Intraspecific variation in clutch size and maternal investment in pueriparous and larviparous Salamandra salamandra females". Evolutionary Ecology (2014): http://www.scopus.com/inward/record.url?eid=2-s2.0-84904273111&partnerID=MN8TOARS.
  55. Azor, J.S.; Santos, X.; Pleguezuelos, J.M.. "Conifer-plantation thinning restores reptile biodiversity in Mediterranean landscapes". Forest Ecology and Management (2014): http://www.scopus.com/inward/record.url?eid=2-s2.0-84931035542&partnerID=MN8TOARS.
  56. Ontiveros, D.; Márquez-Ferrando, R.; Fernández-Cardenete, J.R.; Santos, X.; Caro, J.; Pleguezuelos, J.M.. "Recovery of the Bird Community after a Mine Spill and Landscape Restoration of a Mediterranean River". Restoration Ecology 21 2 (2013): 193-199. http://www.scopus.com/inward/record.url?eid=2-s2.0-84874759778&partnerID=MN8TOARS.
  57. Ballouard, J.-M.; Ajtic, R.; Balint, H.; Brito, J.C.; Crnobrnja-Isailovic, J.; Desmonts, D.; Elmouden, E.H.; et al. "Schoolchildren and one of the most unpopular animals: Are they ready to protect snakes?". Anthrozoos 26 1 (2013): 93-109. http://www.scopus.com/inward/record.url?eid=2-s2.0-84873557155&partnerID=MN8TOARS.
  58. Pleguezuelos, J.M.; Feriche, M.; Santos, X.. "Tail-breakage effects on snake-body condition". Zoologischer Anzeiger 252 2 (2013): 243-245. http://www.scopus.com/inward/record.url?eid=2-s2.0-84875484052&partnerID=MN8TOARS.
  59. Santos, X.; Cheylan, M.. "Taxonomic and functional response of a Mediterranean reptile assemblage to a repeated fire regime". Biological Conservation 168 (2013): 90-98. http://www.scopus.com/inward/record.url?eid=2-s2.0-84885829061&partnerID=MN8TOARS.
  60. Llorente, G.A.; Vidal-García, M.; Garriga, N.; Carranza, S.; Pleguezuelos, J.M.; Santos, X.. "Lessons from a complex biogeographical scenario: Morphological characters match mitochondrial lineages within Iberian Coronella austriaca (Reptilia: Colubridae)". Biological Journal of the Linnean Society 106 1 (2012): 210-223. http://www.scopus.com/inward/record.url?eid=2-s2.0-84859371597&partnerID=MN8TOARS.
  61. Vasconcelos, R.; Santos, X.; Carretero, M.A.. "High temperatures constrain microhabitat selection and activity patterns of the insular Cape Verde wall gecko". Journal of Arid Environments 81 (2012): 18-25. http://www.scopus.com/inward/record.url?eid=2-s2.0-84857995328&partnerID=MN8TOARS.
  62. Santos, X.; Rato, C.; Carranza, S.; Carretero, .M.A.; Pleguezuelos, .J.M.. "Complex phylogeography in the Southern Smooth Snake (Coronella girondica) supported by mtDNA sequences". Journal of Zoological Systematics and Evolutionary Research 50 3 (2012): 210-219. http://www.scopus.com/inward/record.url?eid=2-s2.0-84863553976&partnerID=MN8TOARS.
  63. Garriga, N.; Santos, X.; Montori, A.; Richter-Boix, A.; Franch, M.; Llorente, G.A.. "Are protected areas truly protected? The impact of road traffic on vertebrate fauna". Biodiversity and Conservation 21 11 (2012): 2761-2774. http://www.scopus.com/inward/record.url?eid=2-s2.0-84866119614&partnerID=MN8TOARS.
  64. Santos, X.; Bros, V.; Ros, E.. "Contrasting responses of two xerophilous land snails to fire and natural reforestation". Contributions to Zoology 81 3 (2012): 167-180. http://www.scopus.com/inward/record.url?eid=2-s2.0-84867467132&partnerID=MN8TOARS.
  65. Velo-Antón, G.; Godinho, R.; Harris, D.J.; Santos, X.; Martínez-Freiria, F.; Fahd, S.; Larbes, S.; Pleguezuelos, J.M.; Brito, J.C.. "Deep evolutionary lineages in a Western Mediterranean snake (Vipera latastei/monticola group) and high genetic structuring in Southern Iberian populations". Molecular Phylogenetics and Evolution 65 3 (2012): 965-973. http://www.scopus.com/inward/record.url?eid=2-s2.0-84867738013&partnerID=MN8TOARS.
  66. Montori, A.; Richter-Boix, A.; Franch, M.; Santos, X.; Garriga, N.; Llorente, G.A.. "Natural fluctuations in a stream dwelling newt as a result of extreme rainfall: A 21-year survey of a Calotriton asper population | Fluctuaciones naturales en un tritón de arroyo como resultado de la lluvia extrema: Seguimiento de una población de Calotriton asper durante 21 años". Basic and Applied Herpetology 26 (2012): 43-56. http://www.scopus.com/inward/record.url?eid=2-s2.0-84872131500&partnerID=MN8TOARS.
  67. Santos, X.; Rato, C.; Carranza, S.; Carretero, M.A.; de la Vega, J.P.G.; Pleguezuelos, J.M.. "Morphology matches mtDNA lineages in the southern smooth snake (Coronella girondica) from Iberia". Zoology 115 6 (2012): 398-404. http://www.scopus.com/inward/record.url?eid=2-s2.0-84870255205&partnerID=MN8TOARS.
  68. Mateos, E.; Santos, X.; Pujade-Villar, J.. "Taxonomic and functional responses to fire and post-fire management of a Mediterranean Hymenoptera community". Environmental Management 48 5 (2011): 1000-1012. http://www.scopus.com/inward/record.url?eid=2-s2.0-80054767294&partnerID=MN8TOARS.
  69. Reguera, S.; Santos, X.; Feriche, M.; Mociño-Deloya, E.; Setser, K.; Pleguezuelos, J.M.. "Diet and energetic constraints of an earthworm specialist, the Mesa Central Blotched Garter Snake (Thamnophis scaliger)". Canadian Journal of Zoology 89 12 (2011): 1178-1187. http://www.scopus.com/inward/record.url?eid=2-s2.0-82555194596&partnerID=MN8TOARS.
  70. Bros, V.; Moreno-Rueda, G.; Santos, X.. "Does postfire management affect the recovery of Mediterranean communities? The case study of terrestrial gastropods". Forest Ecology and Management 261 3 (2011): 611-619. http://www.scopus.com/inward/record.url?eid=2-s2.0-78651260009&partnerID=MN8TOARS.
  71. Brito, J.C.; Fahd, S.; Martínez-Freiría, F.; Tarroso, P.; Larbes, S.; Pleguezuelos, J.M.; Santos, X.. "Climate change and peripheral populations: Predictions for a relict Mediterranean viper". Acta Herpetologica 6 1 (2011): 105-118. http://www.scopus.com/inward/record.url?eid=2-s2.0-79959916790&partnerID=MN8TOARS.
  72. Santos, X.; Feriche, M.; León, R.; Filippakopoulou, A.; Vidal-García, M.; Llorente, G.A.; Pleguezuelos, J.M.. "Tail breakage frequency as an indicator of predation risk for the aquatic snake Natrix maura". Amphibia Reptilia 32 3 (2011): 375-383. http://www.scopus.com/inward/record.url?eid=2-s2.0-80052325537&partnerID=MN8TOARS.
  73. Pleguezuelos, J.M.; Brito, J.C.; Fahd, S.; Feriche, M.; Mateo, J.A.; Moreno-Rueda, G.; Reques, R.; Santos, X.. "Setting conservation priorities for the Moroccan herpetofauna: The utility of regional red lists". ORYX 44 4 (2010): 501-508. http://www.scopus.com/inward/record.url?eid=2-s2.0-78650420543&partnerID=MN8TOARS.
  74. Pleguezuelos, J.M.; Feriche, M.; Reguero, S.; Santos, X.. "Patterns of tail breakage in the ladder snake (Rhinechis scalaris) reflect differential predation pressure according to body size". Zoology 113 5 (2010): 269-274. http://www.scopus.com/inward/record.url?eid=2-s2.0-78149361169&partnerID=MN8TOARS.
  75. Santos, X.; Poquet, J.M.. "Ecological succession and habitat attributes affect the postfire response of a Mediterranean reptile community". European Journal of Wildlife Research 56 6 (2010): 895-905. http://www.scopus.com/inward/record.url?eid=2-s2.0-78649347379&partnerID=MN8TOARS.
  76. Márquez-Ferrando, R.; Santos, X.; Pleguezuelos, J.M.; Ontiveros, D.. "Bioaccumulation of heavy metals in the lizard psammodromus algirus after a tailing-dam collapse in Aznalcóllar (Southwest Spain)". Archives of Environmental Contamination and Toxicology 56 2 (2009): 276-285. http://www.scopus.com/inward/record.url?eid=2-s2.0-58149478131&partnerID=MN8TOARS.
  77. Santos, X.; Brito, J.C.; Caro, J.; Abril, A.J.; Lorenzo, M.; Sillero, N.; Pleguezuelos, J.M.. "Habitat suitability, threats and conservation of isolated populations of the smooth snake (Coronella austriaca) in the southern Iberian Peninsula". Biological Conservation 142 2 (2009): 344-352. http://www.scopus.com/inward/record.url?eid=2-s2.0-58149465000&partnerID=MN8TOARS.
  78. Ribeiro, R.; Santos, X.; Sillero, N.; Carretero, M.A.; Llorente, G.A.. "Biodiversity and Land uses at a regional scale: Is agriculture the biggest threat for reptile assemblages?". Acta Oecologica 35 2 (2009): 327-334. http://www.scopus.com/inward/record.url?eid=2-s2.0-60149112127&partnerID=MN8TOARS.
  79. Márquez-Ferrando, R.; Pleguezuelos, J.M.; Santos, X.; Ontiveros, D.; Fernández-Cardenete, J.R.. "Recovering the reptile community after the mine-tailing accident of aznalc.óllar (Southwestern Spain)". Restoration Ecology 17 5 (2009): 660-667. http://www.scopus.com/inward/record.url?eid=2-s2.0-70349512131&partnerID=MN8TOARS.
  80. Santos, X.; Bros, V.; Miño, À.. "Recolonization of a burned Mediterranean area by terrestrial gastropods". Biodiversity and Conservation 18 12 (2009): 3153-3165. http://www.scopus.com/inward/record.url?eid=2-s2.0-70350400452&partnerID=MN8TOARS.
  81. Martínez-Freiría, F.; Santos, X.; Pleguezuelos, J.M.; Lizana, M.; Brito, J.C.. "Geographical patterns of morphological variation and environmental correlates in contact zones: A multi-scale approach using two Mediterranean vipers (Serpentes)". Journal of Zoological Systematics and Evolutionary Research 47 4 (2009): 357-367. http://www.scopus.com/inward/record.url?eid=2-s2.0-70350516755&partnerID=MN8TOARS.
  82. Santos, X.; Llorente, G.A.. "Decline of a common reptile: Case study of the viperine snake Natrix maura in a Mediterranean wetland". Acta Herpetologica 4 2 (2009): 161-169. http://www.scopus.com/inward/record.url?eid=2-s2.0-77957122173&partnerID=MN8TOARS.
  83. Santos, X.; Llorente, G.A.. "Gastrointestinal responses to feeding in a frequently feeding colubrid snake (Natrix maura)". Comparative Biochemistry and Physiology - A Molecular and Integrative Physiology 150 1 (2008): 75-79. http://www.scopus.com/inward/record.url?eid=2-s2.0-43049120955&partnerID=MN8TOARS.
  84. Feriche, M.; Pleguezuelos, J.M.; Santos, X.. "Reproductive ecology of the Montpellier snake, Malpolon monspessulanus (Colubridae), and comparison with other sympatric colubrids in the Iberian Peninsula". Copeia 2 (2008): 279-285. http://www.scopus.com/inward/record.url?eid=2-s2.0-47049123490&partnerID=MN8TOARS.
  85. Brito, J.C.; Santos, X.; Pleguezuelos, J.M.; Sillero, N.. "Inferring evolutionary scenarios with geostatistics and geographical information systems for the viperid snakes Vipera latastei and Vipera monticola". Biological Journal of the Linnean Society 95 4 (2008): 790-806. http://www.scopus.com/inward/record.url?eid=2-s2.0-56649103490&partnerID=MN8TOARS.
  86. Santos, X.; Pleguezuelos, J.M.; Brito, J.C.; Llorente, G.A.; Parellada, X.; Fahd, S.. "Prey availability drives geographic dietary differences of a Mediterranean predator, the Lataste's viper (Vipera latastei)". Herpetological Journal 18 1 (2008): 16-22. http://www.scopus.com/inward/record.url?eid=2-s2.0-50249131498&partnerID=MN8TOARS.
  87. Santos, X.; Roca, J.; Pleguezuelos, J.M.; Donaire, D.; Carranza, S.. "Biogeography and evolution of the Smooth snake Coronella austriaca (Serpentes: Colubridae) in the Iberian Peninsula: Evidence for Messinian refuges and Pleistocenic range expansions". Amphibia Reptilia 29 1 (2008): 35-47. http://www.scopus.com/inward/record.url?eid=2-s2.0-56649096988&partnerID=MN8TOARS.
  88. Santos, X.; Llorente, G.A.; Pleguezuelos, J.M.; Brito, J.C.; Fahd, S.; Parellada, X.. "Variation in the diet of the Lataste's viper Vipera latastei in the Iberian Peninsula: Seasonal, sexual and size-related effects". Animal Biology 57 1 (2007): 49-61. http://www.scopus.com/inward/record.url?eid=2-s2.0-33947614645&partnerID=MN8TOARS.
  89. Santos, X.; Brito, J.C.; Pleguezuelos, J.M.; Llorente, G.A.. "Comparing Filippi and Luiselli's (2000) method with a cartographic approach to assess the conservation status of secretive species: The case of the Iberian snake-fauna". Amphibia Reptilia 28 1 (2007): 17-23. http://www.scopus.com/inward/record.url?eid=2-s2.0-34547946576&partnerID=MN8TOARS.
  90. Pleguezuelos, J.M.; Santos, X.; Brito, J.C.; Parellada, X.; Llorente, G.A.; Fahd, S.. "Reproductive ecology of Vipera latastei, in the Iberian Peninsula: Implications for the conservation of a Mediterranean viper". Zoology 110 1 (2007): 9-19. http://www.scopus.com/inward/record.url?eid=2-s2.0-33845878536&partnerID=MN8TOARS.
  91. Santos, X.; Arenas, C.; Llorente, G.A.; Ruiz, X.. "Exploring the origin of egg protein in an oviparous water snake (Natrix maura)". Comparative Biochemistry and Physiology - A Molecular and Integrative Physiology 147 1 (2007): 165-172. http://www.scopus.com/inward/record.url?eid=2-s2.0-33947540908&partnerID=MN8TOARS.
  92. Santos, X.; Llorente, G.A.; Montori, A.; Carretero, M.A.; Franch, M.; Garriga, N.; Richter-Boix, A.. "Evaluating factors affecting amphibian mortality on roads: The case of the Common Toad Bufo bufo, near a breeding place". Animal Biodiversity and Conservation 30 1 (2007): 97-104. http://www.scopus.com/inward/record.url?eid=2-s2.0-35448934471&partnerID=MN8TOARS.
  93. Segura, C.; Feriche, M.; Pleguezuelos, J.M.; Santos, X.. "Specialist and generalist species in habitat use: Implications for conservation assessment in snakes". Journal of Natural History 41 41-44 (2007): 2765-2774. http://www.scopus.com/inward/record.url?eid=2-s2.0-38849131436&partnerID=MN8TOARS.
  94. Brito, J.C.; Santos, X.; Pleguezuelos, J.M.; Fahd, S.; Llorente, G.A.; Parellada, X.. "Morphological variability of the Lataste's viper (Vipera latastei) and the Atlas dwarf viper (Vipera monticola): Patterns of biogeographical distribution and taxonomy". Amphibia Reptilia 27 2 (2006): 219-240. http://www.scopus.com/inward/record.url?eid=2-s2.0-33645422521&partnerID=MN8TOARS.
  95. Santos, X.; Martínez-Freiría, F.; Pleguezuelos, J.M.; Roca, V.. "First helminthological data on Iberian vipers: Helminth communities and host-parasite relationships". Acta Parasitologica 51 2 (2006): 130-135. http://www.scopus.com/inward/record.url?eid=2-s2.0-33745870124&partnerID=MN8TOARS.
  96. Santos, X.; Brito, J.C.; Sillero, N.; Pleguezuelos, J.M.; Llorente, G.A.; Fahd, S.; Parellada, X.. "Inferring habitat-suitability areas with ecological modelling techniques and GIS: A contribution to assess the conservation status of Vipera latastei". Biological Conservation 130 3 (2006): 416-425. http://www.scopus.com/inward/record.url?eid=2-s2.0-33645383995&partnerID=MN8TOARS.
  97. Santos, X.; Vilardebó, E.; Casals, F.; Llorente, G.A.; Vinyoles, D.; Sostoa, A.. "Wide food availability favours intraspecific trophic segregation in predators: The case of a water snake in a Mediterranean river". Animal Biology 56 3 (2006): 299-309. http://www.scopus.com/inward/record.url?eid=2-s2.0-33749508884&partnerID=MN8TOARS.
  98. Montori, A.; De Figueroa, J.M.T.; Santos, X.. "The diet of the brown trout Salmo trutta (L.) during the reproductive period: Size-related and sexual effects". International Review of Hydrobiology 91 5 (2006): 438-450. http://www.scopus.com/inward/record.url?eid=2-s2.0-33750713053&partnerID=MN8TOARS.
  99. Santos, X.; Llorente, G.A.; Feriche, M.; Pleguezuelos, J.M.; Casals, F.; de Sostoa, A.. "Food availability induces geographic variation in reproductive timing of an aquatic oviparous snake (Natrix maura)". Amphibia Reptilia 26 2 (2005): 183-191. http://www.scopus.com/inward/record.url?eid=2-s2.0-23444433200&partnerID=MN8TOARS.
  100. Martin, J.E.; Llorente, G.A.; Roca, V.; Carretero, M.A.; Montori, A.; Santos, X.; Romeu, R.. "Relationship between diet and helminths in Gallotia caesaris (Sauria: Lacertidae)". Zoology 108 2 (2005): 121-130. http://www.scopus.com/inward/record.url?eid=2-s2.0-20444392809&partnerID=MN8TOARS.
  101. Santos, X.; Llorente, G.A.. "Lipid dynamics in the viperine snake, Natrix maura, from the Ebro Delta (NE Spain)". Oikos 105 1 (2004): 132-140. http://www.scopus.com/inward/record.url?eid=2-s2.0-1542438839&partnerID=MN8TOARS.
  102. Santos, X.; Llorente, G.A.. "Seasonal variation in reproductive traits of the oviparous water snake, Natrix maura, in the Ebro Delta of northeastern Spain". Journal of Herpetology 35 4 (2001): 653-660. http://www.scopus.com/inward/record.url?eid=2-s2.0-0035660903&partnerID=MN8TOARS.
  103. Santos, X.; González-Solís, J.; Llorente, G.A.. "Variation in the diet of the viperine snake Natrix maura in relation to prey availability". Ecography 23 2 (2000): 185-192. http://www.scopus.com/inward/record.url?eid=2-s2.0-0342424369&partnerID=MN8TOARS.
  104. Santos, X.; Pastor, D.; Llorente, G.A.; Albaigés, J.. "Organochlorine levels in Viperine Snake Natrix maura carcasses from the Ebro Delta (NE Spain): Sexual and size-related differences". Chemosphere 39 15 (1999): 2641-2650. http://www.scopus.com/inward/record.url?eid=2-s2.0-0033485414&partnerID=MN8TOARS.
  105. Santos, X.; Llorente, G.A.. "Sexual and size-related differences in the diet of the snake Natrix maura from the Ebro Delta, Spain". Herpetological Journal 8 3 (1998): 161-165. http://www.scopus.com/inward/record.url?eid=2-s2.0-0031752260&partnerID=MN8TOARS.


Título / Tema
Papel desempenhado
Curso (Tipo)
Instituição / Organização
2016 - 2019/04/15 Analysis of the resilience to fire of the reptile community and the forest in the western Rif Mountains (northern Morocco)
University of Abdelmalek Eassaadi, Marrocos
2012 - 2016/01/20 Ecología trófica de tres especies de serpientes de cascabel en México: Crotalus aquilus, Crotalus polystictus y Crotalus willardi.
Universidad de Granada Departamento de Zoología, Espanha
2015 - 2016 Stable isotope analyses of Invertebrate taxa from the Island of Socotra (Yemen) Master of Biodiversity
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2014 - 2015 Uncovering the phylogeography of the habitat-specialist land snail Xerocrassa montserratensis
Universitat de Barcelona Departament de Genètica Microbiologia i Estadística, Espanha
2013 - 2015 Genetic and morphological impacts of a repeated fire regime on Podarcis guadarramae
Universidade do Porto Centro de Investigação em Biodiversidade e Recursos Genéticos, Portugal
2013 - 2015 The impact of fire on mammal species: a meta-analysis and the particular case of the Iberian wolf (Canis lupus signatus) in Portugal
Universidade do Porto Centro de Investigação em Biodiversidade e Recursos Genéticos, Portugal
2013 - 2014 Respuesta de la comunidad de reptiles a tratamientos de aclareo de repoblaciones forestales en Sierra Nevada
Universidad de Granada Departamento de Zoología, Espanha
2013 - 2014 The mechanisms that produce speciation: diet description of Socotra reptiles
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2013 - 2014 Mechanisms that produce speciation: Description of stable isotopes of Socotran Reptiles
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2013 - 2014 Diet of Mauritanian population of the west African crocodile (Crocodylus suchus) examined with stable isotopes
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2011 - 2012 Effects of repeated-fire regimes on the Hermann’s tortoise Testudo hermanni and on the reptile communities at the Albera Range (Girona, North-eastern Catalonia)
Aix-Marseille Université, França
2011 - 2012 Estrategias Antidepredatorias en la culebra viperina Natrix maura. Huida, coloración disruptiva y mimetismo batesiano
Universidad de Granada Departamento de Zoología, Espanha
2009 - 2010 Effect of Prey availability on growth rates in Natrix maura at the Iberian Peninsula Máster en Biodiversidad Animal de la Universidad de Barcelona
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2009 - 2010 Dietary and energetic traits of a vermivorous snake, Thamnophis scaliger
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2003 - 2008 Ecology of restored zones: response of the bird and reptile communities in restored areas of Mediterranean habitats
Universidad de Granada Departamento de Zoología, Espanha
2006 - 2007 Land Use and Biodiversity at a Regional Scale: the case of the Catalonian Reptiles
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha
2004 - 2005 Abiotic and biotic parametters which determined the occurrence of three herp species in Catalan Rivers
Universitat de Barcelona Departament de Biologia Evolutiva Ecologia i Ciències Ambientals, Espanha

Outra distinção

2001 Extraordinary Award of PhD
Universitat de Barcelona, Espanha